They are nice clubs.  Speaking as a botanist, I don't think that the heads are intended to represent a fruit or nut, at least one I can recognize.  My guess is that the pieces may have been initially turned on a lathe, and that the indentations in the the tips of the heads might be where the spur from a lathe was used to hold the piece as it was turned.  The longitudinal lines would have been added later.   
 
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